Table 4 The statistical cross-fostering design matrix
CombinationCoefficients
Raodoioamdmcmoddmo
M11O1110Embedded Image00Embedded Image1Embedded Image
M11O2110Embedded Image01Embedded Image0Embedded Image
M12O1111Embedded Image00Embedded Image0Embedded Image
M12O2110Embedded Image−10Embedded Image0Embedded Image
M12O1210Embedded Image10Embedded Image0Embedded Image
M12O221−1Embedded Image00Embedded Image0Embedded Image
M22O1210Embedded Image0−1Embedded Image0Embedded Image
M22O2210Embedded Image00Embedded Image−1Embedded Image
  • The rows are labeled to reflect the corresponding maternal–offspring genotype combination, as in Table 3 [given as MijOkl, where Mij is the unordered genotype of the mother (with i and j being the two alleles she has at the focal locus) and Okl is the ordered genotype of the offspring (with again, k and l being the two alleles that the offspring has at the focal locus]. The columns labeled “Coefficients” correspond to the index scores used in a regression model to fit the intercept or reference point for the model (R), the three direct genetic effects (ao, do, and io), and the two maternal effects (am and dm). The last two columns correspond to terms that we do not interpret here because they do not correspond to independent effects of the maternal and offspring genomes. The first of these, cmo, is the confounded effect (see text) and the second, dd, is an interaction between the dominance effects in mothers and their offspring (see Discussion)