f, F | Distribution of final gap δ_{f} |

K | Carrying capacity |

L | No. of loci |

p | Distribution of phenotypic effects of new mutations |

q | Distribution of fitness effects of new mutations |

R | Transition density for distribution of initial gap ρ(δ_{i}) |

s | Selection coefficient |

t | Time |

u | Mutation rate per locus |

v | Speed of the optimum |

w | Fitness |

z | Phenotype |

z_{opt} | Optimal phenotype at a given time |

z_{max} | Maximal value of z_{opt} in some simulations |

α | Phenotypic effect of mutation (step size) |

γ | Composite parameter relating ecological and genetic factors (v/[σθ]) |

δ | Phenotypic gap (z – z_{opt}) |

δ_{i} | “Initial” phenotypic gap before an adaptive step |

δ_{f} | “Final” phenotypic gap before an adaptive step |

Θ | Population and trait-wide mutation rate (2KLu) |

μ | Mean effect of positive new mutations |

ρ | Distribution of initial gap δ_{i} |

σ | Strength of selection |

ϕ | Distribution of phenotypic effects of adaptive substitutions |

χ | Distribution of fitness effects of adaptive substitutions |

Ψ | “Selective sieve” |

Ψ_{s} | Static part of selective sieve |

Ψ_{d} | Dynamic part of selective sieve |

ω | Standard deviation of distribution of new mutations p(α) |

α and

*v*are generally measured in units of ω, and σ is measured in units of 1/ω^{2}. The unscaled counterparts are marked by a tilde (*e.g*., ).