Commonly used notation

NTotal population size
μWild-type mutation rate per genome
μ+Mutator mutation rate per genome
UMutation rate into mutator state
LLength of genome
bNumber of 1's in genome
δFraction of mutations that are lethal
xMutator frequency
Embedded ImageAverage mutation rate per genome
R ≡ μ+Mutator strength
rb/LGrowth rate per individual per simulation time step
s = 1/bSelection coefficient of nonlethal mutation
α ≡ 1 − b/LFraction of 0's in the genome
αe ≡ α(1 − δ)Fraction of mutations that are beneficial