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is the genotypic mean of the backcross progenies from F

_{2}parents with marker genotype*g*backcrossed to parental line*j*(*j*= 2, 1). Only coefficients of , , and are given for means. The coefficients of , , and for are the same as those for . , , and have the same coefficients as , , and for ; but for , the coefficients have opposite signs.