Statistic^{a} | ||||
---|---|---|---|---|

Variable X | ε(X) | ε(X | QTL i) | ε(X | QTL i × QTL j) | |

Z_{1} | aa_{ij} + dd_{ij} | |||

Z_{2} | ad_{ij} + da_{ij} | |||

Z_{3} | 2dd_{ij} | |||

H_{3} | μ | a_{i} | aa_{ij} |

For description of

*Z*_{1},*Z*_{2},*Z*_{3}, and*H*_{3}and for notation of genetic effects see materials and methods.↵

*a*Summation is over the entire set of QTL involved in the trait expression.