Symbol | Description |
---|---|

b_{i} | Mutational bias at locus i |

b̃ | Weighted average of mutational biases over all loci |

D(β) | The nonmonotonic function characterizing the mean phenotypic value |

ε,V_{e} | Environmental effect and its variance |

f_{i} | Distribution of mutant allelic effects at locus i |

ϕ(_{i}x)_{i} | Distribution of allelic effects of maternal origin in offspring at locus i |

γ | Mutational regression parameter |

G, G¯ | Genotypic value and its mean equilibrium value |

L | Genetic load |

m_{i} | Standard deviation of mutant allelic effects at locus i |

μ_{i} | Allelic mutation rate at locus i |

n | Number of loci affecting the trait |

s | Strength of selection on genotypic values |

x_{i} | Allelic effect of maternal origin at locus i |

x¯_{i} | Mean allelic effect of maternal origin in offspring at locus i |

x, Φ(x) | Vector of n allelic effects in gametes and its equilibrium distribution |

U | Expected number of new mutations, per individual, per generation |

V_{G} | Variance in genotypic values (genetic variance) |

w(G) | Proportional to the viability of offspring with genotypic value G |

z, z¯ | Phenotypic value and its mean equilibrium value in offspring |

z_{opt} | Optimal phenotypic value |