TABLE 4

Quantitative trait loci for headless behaviors

		Female		Male		Total
Trait (95% LR)^a	Interval	LR	Effect^b	LR	Effect	LR	Effect	Model^cP value
Hindgrooming (14.7)	3E	20.4	-0.48	n.s.		11.6	-0.30	0.0002
	98A	15.5	-0.48	21.0	-0.71	25.9	-0.60	<0.0001
Quivering (37.0)	14C	36.5	-1.63	20.2	-1.00	23.7	-0.58	<0.0001
	87B	n.s.		n.s.		12.5	0.20	0.0005
	14C × 87B							0.0255
Grasping (15.8)	60E	n.s.		20.9	-0.67	n.s.		n.s.
	61A	18.6	-0.42	n.s.		23.4	-0.73	0.0016
	63-65	18.9	0.49	16.3	0.49	n.s.		0.0016
	87B	n.s.		n.s.		16.4	0.55	<0.0001
	60E × 87B							0.0065
Self-righting (14.8)	87B	12.8	0.37	28.0	0.83	18.4	0.44	<0.0001
	96A	13.1	0.50	n.s.		18.9	0.51	0.0009

n.s., not significant.
↵^a 95% significance threshold likelihood ratio (LR) for combined sexes for indicated traits.
↵^b Estimated additive effect of QTL as a fraction of the standard deviation among line means.
↵^c See text for description of two- or three-locus linear multiple regression model.