t,u  Time variables 
n_{c}, q  Number of breeding categories, indexed by q 
m, f  Denotes the two sexes in discrete generations, i.e., q = m (male) or f (female) 
r_{i},r_{i(q)}  Observed longterm genetic contribution of individual i (in category q) 
r_{i,u}(q, t)  The genetic contribution of individual i born at time u to selected parents of sex q born at time t 
F_{t}, ΔF  Inbreeding coefficient at time t, and rate of inbreeding 
ω  Deviation from HardyWeinberg equilibrium 
X_{q}, X  Number of parents in category q and a simple monoecious population, respectively 
C_{u}(t)  Sum of squared contributions for individuals born at time u to selected parents at time t 
C  Converged sum of squared contributions, independent of time in an equilibrium 
L  Generation interval 
S_{i(q)}  Set of selective advantages for individual i in category q 
μ_{i(q)}  Expected contribution of i in category q conditional upon s_{i(q)} 
 Variance of contribution of i in category q conditional upon s_{i(q)} 
n_{i}  Number of selected offspring of i 
θ_{n,i}  Expected number of selected offspring of i conditional upon s_{i}(_{q}) 
V_{n,i}  Variance of the number of selected offspring of i, conditional upon s_{i}(_{q)} 
V_{n,dev,i}  Deviation of V_{n, i} from Poisson, i.e., V_{n,dev,i} = V_{n,i} — θ_{n, i} 
α_{q}, β_{q}  Linear model for
