TABLE 3

Putative QTL positions, LOD thresholds, LOD scores, and magnitude of effects for 15 morphological traits and pollen viability in a backcross population of H. annuus (CMS89) × H. debilis ssp. cucumerifolius

Trait^a	LOD threshold	Linkage group	Position^b	Interval markers^c	LOD score	PVE (%)^d
Speckled internode	2.68	3	13.41	GCCC.209-GCGG.317	3.09	6.3
		8	107.67	CGGG.110-GCCT.164	14.82	27.1
		11	38.34	CGCG.380-CGTG.097	22.26	36.5
Tooth height	2.68	2	28.92	GCCT.245-CGCT.385	11.65	21.9
		2	56.89	CTCT.218-CGGG.054	9.05	32.1
		5	33.28	GCGC.290-CTAT.191	6.41	13.8
Head no.	2.80	6	27.70	GCCC.108-CGCT.048	7.06	13.4
		7	131.78	GCCC.175-GCTC.150	5.11	9.9
Ray no.	2.69	2	73.89	CGCG.086-GCGC.136	4.21	8.7
		3	24.99	GCTC.210-CGTG.116	3.46	9.1
		5	8.01	CTCT.211-GCGG.293	4.13	8.8
		8	115.96	CGGG.235-GCGG.110	5.79	11.8
Ligule length	2.72	1	10.01	CGGG.050-CGTG.055	6.08	16.2
		5	37.28	GCGC.290-CTAT.191	3.91	8.9
		7	131.78	GCCC.175-GCTC.150	3.70	7.6
Ligule width	2.72	2	43.84	CTCT.209-CTCT.218	6.10	12.9
		6	27.70	GCCC.108-CGCT.048	2.81	5.8
		9	0.01	CGCG.360-CGTG.226	4.41	9.0
Ligule shape	2.75	1	52.49	CGTG.308-CGCT.135	3.36	6.9
		2	81.49	GCGA.140-GCCT.199	17.83	32.7
		6	26.72	GCAG.066-GCCC.108	3.87	7.9
		9	0.01	CGCG.360-CGTG.226	3.86	7.9
		13	0.01	CTCT.063-CTCT.133	3.25	6.7
Disk diameter	2.63	6	29.26	CGCT.048-GCGG.151	3.59	7.1
		8	105.67	CGGG.110-GCCT.164	4.93	10.2
		8	117.96	CGGG.235-GCGG.110	5.04	10.5
Phyllary length	2.53	4	29.60	CGCT.169-GCCC.178	2.88	6.8
		5	33.28	GCGC.290-CTAT.191	4.14	10.8
		11	49.68	GCCC.050-GCCC.071	5.40	12.1
Phyllary width	2.66	3	44.29	GCGC.125-GCCT.101	5.20	10.7
		4	72.01	CTAT.306-GCGG.152	3.20	9.4
		6	24.59	CGCT.241-GCAG.066	2.85	6.5
		7	131.78	GCCC.175-GCTC.150	3.72	7.7
		12	15.69	CTCT.127-CTCT.116	3.12	6.6
Phyllary shape	2.65	3	44.29	GCGC.125-GCCT.101	4.42	9.3
		4	9.64	CGCT.353-CGCT.169	2.79	5.9
		6	27.70	GCCC.108-CGCT.048	4.08	8.4
		8	103.67	CGGG.110-GCCT.164	3.57	7.4
Phyllary pubescence	2.66	4	64.63	CGCG.124-GCGA.184	5.40	10.4
		5	23.25	CGCT.234-CGCT.322	10.18	20.6
		6	33.24	GCCT.048-CGGG.045	8.78	16.3
		8	108.92	GCCT.164-GCAG.094	4.01	7.8
Achene length	2.25	1	8.01	CGGG.050-CGTG.055	6.98	16.4
		1	63.34	GCTC.163-CGGG.205	8.79	18.1
		1	86.59	GCCT.295-GCGG.171	7.54	15.2
		2	77.22	GCCT.156-GCGA.140	2.56	5.4
Achene width	2.65	2	33.99	GCTC.068-GCGC.099	3.53	7.5
		3	16.53	GCGG.317-GCTC.210	5.46	11.2
		6	26.72	GCAG.066-GCCC.108	8.56	17.1
		7	131.78	GCCC.175-GCTC.150	2.78	5.9
Achene shape	2.66	1	38.59	CGTG.055-CGTG.308	5.47	13.2
		1	71.51	CGGG.205-CGCT.163	4.90	11.2
		2	19.76	GCGG.065-CTAT.060	3.54	8.1
		5	19.25	CGCT.234-CGCT.322	3.56	7.6
		6	27.70	GCCC.108-CGCT.048	2.68	5.7
		8	22.01	GCCC.044-GCTC.083	5.86	19.5
Pollen viability	3.98	2	27.44	GCGG.065-CTAT.060	7.47	38.2
		8	38.00	GCTC.083-GCGA.295	4.57	38.8

↵^a Nonintrogressed traits are in italics.
↵^b Absolute position from left telomere in cM.
↵^c Marker nearest to QTL is underlined.
↵^d Proportion of phenotypic variation explained by QTL using CIM. QTL effects in the direction of the H. annuus phenotypes are in italic type.