Parameter | Identifier | Mean | Standard deviation |
---|---|---|---|

Base frequency of A | π_{A} | 32.7% | — |

Base frequency of C | π_{C} | 33.8% | — |

Base frequency of G | π_{G} | 11.4% | — |

Base frequency of T | π_{T} | 22.1% | — |

Transition/transversion | κ | 15.5 | 5.0 |

Pyrimidine/purine trans. | ξ | 1.85 | 0.48 |

Rate heterogeneity | α | 0.26 | 0.05 |