Drift Measure | F_{2} (P_{1}, P_{2}) | F_{3} (P; _{X}P_{1}, P_{2}) | F_{4}(P_{1}, P_{2}, P_{3}, P_{4}) |
---|---|---|---|

Definition | |||

F_{2} | — | ||

Coalescence times | |||

Variance | |||

Branch length |

A constant of proportionality is omitted for coalescence times and branch lengths. Derivations for

*F*_{2}are given in the main text, and*F*_{3}and*F*_{4}are a simple result of combining Equation 16 with Equations 20b and 24b. and correspond to the average length of the internal branch in a gene genealogy concordant and discordant with the population assignment, respectively (see*Gene tree branch lengths*section).