Table 1 Summary of equations
Drift MeasureF2 (P1, P2)F3 (PX; P1, P2)F4(P1, P2, P3, P4)
DefinitionEmbedded ImageEmbedded ImageEmbedded Image
F2Embedded ImageEmbedded Image
Coalescence timesEmbedded ImageEmbedded ImageEmbedded Image
VarianceEmbedded ImageEmbedded ImageEmbedded Image
Branch lengthEmbedded ImageEmbedded ImageEmbedded Image
  • A constant of proportionality is omitted for coalescence times and branch lengths. Derivations for F2 are given in the main text, and F3 and F4 are a simple result of combining Equation 16 with Equations 20b and 24b. Embedded Image and Embedded Image correspond to the average length of the internal branch in a gene genealogy concordant and discordant with the population assignment, respectively (see Gene tree branch lengths section).