vol. 192 no. 3 973-985
- Received June 29, 2012.
- Accepted July 27, 2012.
- Published online November 7, 2012.
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Copyright © 2012 by the Genetics Society of America
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- *Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania 19104
- †Cell Biology and Molecular Genetics, Center for Bioinformatics and Computational Biology, University of Maryland, College Park, MD 20742
- ↵1Corresponding author: Department of Biology, University of Pennsylvania, Philadelphia, PA 19104. E-mail: