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Corresponding author: Dominique Mouchiroud, Centre National de la Recherche Scientifique UMR 5558, Biométrie, Génétique et Biologie des Populations, Université Claude Bernard Lyon 1, 43, Boulevard du 11 novembre 1918, 69622 Villeurbanne Cedex, France., mouchi{at}biomserv.univ-lyon1.fr (E-mail).
Communicating editor: G. A. CHURCHILL
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